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Predicting tree heights from a
combination of LIDAR canopy heights and digital stem counts
S. Magnussen, F. Gougeon, D. Leckie and M. Wulder
Natural Resources Canada, Canadian Forestry Service,
Pacific Forestry Center, 506 W. Burnside Road, Victoria, British
Columbia, Canada, V8Z 1M5.
Email: {smagnuss| fgougeon| dleckie| mwulder
@pfc.forestry.ca}
Abstract
Tree heights can be derived from LIDAR (Light
Detection And Ranging) data of canopy heights when an estimate of the
expected number (N) of ‘tree top’ hits exists. The probability of
a LIDAR pulse hitting a tree top is governed by the number and size of
‘tree tops’. Counts of trees with a sunlit crown were derived from
Compact Airborne Spectrographic Imager images with 55 x 55 cm
pixels. Pairing this count with an assumed ‘tree top’ size of 3 pixels
suggested an average of 6 ‘tree top’ hits in a 400 m2
plot. The topmost Nhit canopy heights can be considered as
proxies of tree height. This simple concept is demonstrated in a thinning
trial with 39-year-old Douglas-fir. Extreme value distributions fitted to
expected ‘tree-top’ hits generated various statistics of predicted
dominant and codominant tree heights. Our approach lowered the average
discrepancy between mean canopy height and Lorey’s mean tree height from
20% to an insignificant 3% (P=0.09).
Ground-based tree-counts improved the results slightly.
1. Introduction
Accurate canopy height profiling of forest stands is now a reality due to improvements of differential
global positioning systems, laser scanner technology, and inertial navigation systems (Flood and Gutelius 1997, Ritchie
1995, Blair et al. 1994, Bufton 1989). Canopy height data provide valuable information about the forest stand, canopy
structure and possibly crown closure (Nelson 1997, Ritchie et al. 1993). However, to estimate mean stem volume and site
productivity of a stand a measure of stand height is needed. Canopy height is the vertical distance from an arbitrary
point on the forest floor to the topmost part of the canopy at the chosen location. Tree height is the vertical distance
from the base of a tree to its topmost terminal bud (Avery and Burkhart 1983). The relationship between the two can be
very complex. For dominant and codominant trees the mean canopy height is bound to be lower than the mean tree height
(Avery and Burkhart 1983). Crown shape, stem density and spatial distribution of the trees will all influence the
discrepancy.
Laser scanning of forest stands provides the mean canopy height of dominant and codominant trees. The
chance that a laser pulse hits an intermediate or even a suppressed tree is low (Magnussen and Boudewyn 1998). Converting
canopy heights to tree heights can be done by either a model-based approach (Magnussen and Boudewyn 1998) or through
heuristic methods consisting of selecting the largest canopy height within spatially binned areas (N¶sset 1997, Nilsson
1996). Extreme values of canopy heights are more likely to be associated with a tree top than less extreme values. Both
approaches have demonstrated their validity. The advantages of a model-based approach include a reduced reliance on
auxiliary information from ground-based surveys. Model-based approaches, on the other hand, tend to be complex and
computationally demanding (Magnussen et al. 1999). This study demonstrates how tree counts and an estimate of the
combined area occupied by tree "tops” (target area) leads to improved estimates of tree heights from laser canopy height
data.
2. Material and Methods
Study site
Data were collected from one 50-ha stand of 49-year-old second-growth Douglas-fir near Shawnigan Lake ( 48o 38' N, 123o 43' W) on Vancouver Island of British Columbia. The stand was fairly homogenous and on a gentle slope. The stand contained a thinning and fertilization trial (Brix 1993) with 42 square plots of 400 m2 each. Geographic coordinates for each plot were obtained by bearing and distance from a known control point.
Field data
All trees in each plot were measured in the
winter of 1994-1995 (age 49) for diameter at breast height (DBH cm, 1.3 m
above ground). At that time the plots contained between 32 and 158 trees
per plot (median 64, mean 66, standard deviation 31.9). Fifteen trees per
plot representing the plot range of DBH values were also measured for
height (HT). HT of the remaining trees were predicted from plot-specific
nonlinear height diameter relationships (McWilliams and Therien
1997). DBH and HT values of all trees were prorated to their expected
1995-1996 status by means of individual and trait-specific relative
growth rates (RGR). The RGR values used were those registered for the
period between the last two measurements (1989/90 and 1994/95). A control
sample of 46 trees was established to check the predicted height
growth. A t-test of predicted versus measured heights did not reject the
hypothesis of no difference (P=0.29). Heights of dominant and codominant
trees were computed as Lorey’s height (Löetch and Haller 1964), i.e. the
basal-area weighted tree height.
Crown shape data were obtained from a 1992 biomass
study of 120 trees from 12 plots. The study furnished 2214 paired
measurements of branch length (BL) and the distance (DT) between the stem
location of the branch and the top of the tree (Brix 1993). Branch length
(BL) was found to be a power function of distance to the top (DT). The
average relationship was BL = DT0,421 (root mean square
error = 0.19, r2=0.91). This functional relationship was used
to compute the expected total crown area at various canopy depths.
2.3 CASI data
The Compact Airborne Spectrographic Imager
(CASI) is an airborne sensor capable of acquiring multispectral imagery
in the visible and near infrared part of the spectrum (Anger et
al. 1994). CASI was flown to acquire a high spatial resolution
multispectral image (nine spectral bands, mostly ~14 nm wide) over the
study site. Using an onboard inertial navigation system, differential GPS
technology and proprietary algorithms, the supplier provided the image
already geo-referenced to UTM coordinates and at a nominal spatial
resolution of 55 cm/pixel.
2.4 Digital stem counts on CASI images
Stem counts were obtained from the CASI image using a
technique originally developed to detect and classify the species of
mature trees in medium resolution airborne sensor images (1-2
m/pixel) (Gougeon and Moore 1989). The technique consists of detecting
local maxima in a smoothed image from the most appropriate spectral band
(typically the near infrared). In medium to dense coniferous stands,
where individual tree crowns appear separated by areas of shade, the
algorithm generally isolates a single pixel per tree, usually
corresponding closely to a well illuminated tree top. An appropriately
chosen threshold suppresses local maxima in the shaded areas. Recent
applications of this technique includes regeneration assessment in dense,
moderate, and even open stands using 30 cm/pixel MEIS images (Gougeon,
1997). Similar techniques have been used by Eldrige and Edwards (1993),
and Dralle and Rudemo (1996). In this study we improved the stem count
technique by "doubling the spatial resolution” (to 27.5 cm/pixel) before
smoothing the image.
Laser data
Laser scanner data were acquired in December
1996 with the Optech ALTM 1020 laser scanning system supplied with a
Trimble 4000 SSE GPS receiver. Two flight lines were each flown in a
forward direction then backward recording distances to the first and the
last laser pulse, respectively. Flying altitude was about 600 m with a
ground speed of 40 m·s-1. Laser pulses were emitted at a
wavelength of 1047 nm with a frequency of 2 to 8 kHz. Scan angles were
less than ±12o.
Two data files were generated from the flights and
some proprietary post-flight processing: one containing 25835 triplets of
latitude (X), longitude (Y), and elevation (Z) of first returned pulses
and the second 17413 triplets (X, Y, Z) of last returned pulses. The
former is assumed to constitute ‘canopy hits’ and the latter ‘ground
hits’. We refer to these data as (Xc , Yc ,
Zc) for the imputed canopy hits, and (Xg ,
Yg , Zg) for the imputed ground hits. Positional
errors across the scanned area are 1 to 3 m for x and y and about 20 cm
for z (Magnussen and Boudewyn 1998). That is, all hits from a single
flight line may have a constant location error of 1 to 3 m in either
horizontal direction. Relative positions of points gathered in a single
over-flight are accurate to within about 20 cm. Six of the 42 plots were
only partially scanned by the over-flight, leaving 36 plots for the
analyses. Further details about the data capture are available (Magnussen
and Boudewyn 1998).
With a beam divergence of 0.25 mrad, an average flight height of 600 m, and an average canopy height of 18 m the laser footprint of canopy hits is a circle with radius about 0.14 cm.
Laser canopy heights
Canopy heights were computed as  where is a prediction of the ground elevation at the location (xc , yc) obtained from a digital elevation model (DEM) fitted to the ground triplets (Magnussen and Boudewyn 1998). DEM residuals had a standard error of about 20 cm and appeared to be normally distributed.
Scatter plots of versus indicated a large number of near ground canopy heights. Thus
we conjectured that the canopy hits were a mixture of ‘near ground’ hits
and actual canopy hits. Application of an Estimation-Maximization
(EM) algorithm (Dempster et al. 1977) based on two distinct linear
relationships between and in this mixture enabled us to mark 14.6% of the imputed canopy
hits as ‘near ground’ hits.The ‘near ground’ data were discarded from
the analyses; Magnussen and Boudewyn (1998) provide further
details. Canopy heights (CH) were then equated to for the remaining 22061
records. When allocated to the 36 plots each field plot contained between
29 and 71 estimates of canopy height (median 44, mean 48, standard
deviation 11.4).
2.7 Expected number of laser pulses hitting a tree "top”
An estimate of the number of laser pulses hitting a
tree "top” 
is obtained
from:
(1) 
where 
denote the number of laser pulses hitting the canopy within an area A,

an estimate of the
number of visible tree tops within area A, and the average horizontally exposed
area of a tree top. Exposed in this context simply means that the crown
area is visible from above. Implicit in (1) is the assumption that laser
pulses and trees are randomly distributed within A (Cressie
1991). Note in (1) includes the convolution of the actual target area (tree
top) with the spatial extension of the laser pulses (Stoyan et
al. 1987).
CASI-based estimation of and 
For the CASI-based estimates of derived from the automated stem
detection, a target area of 3 pixels was assumed. While the tree top proper surely
occupies less than one pixel the convolution with the laser footprint and
allowance for hits ‘near’ (within the topmost annual growth whorl) the
top makes the assumption reasonable.
Ground-based estimation of and
Ground-based tree-counts ( ) while readily available need to
be converted before they enter (1). In general, as some trees are obscured from
the view of the laser pulses by adjacent trees. Also, the average target
area 
must be determined from
considerations based on the average geometry of tree crowns.
, in this context, is defined as the horizontal projection of
the space defined by the uppermost whorl and the terminal bud of a tree
(a cone). The projected space is assumed to be circular with a radius
equal to  where is the average current height
increment of the trees (Magnussen and Boudewyn 1998). For a sub-sample
of 46 randomly selected trees we found which gives a target radius of
about 0.6 m. The effective target radius is slightly larger if we allow
half a laser footprint inside a target area to trigger a hit. From these
considerations we arrived at  
1.7 m2.
Conversion of to 
relied on
considerations of canopy geometry. We essentially computed the
probability that a tree would have an exposed tree top (PET
) and then derived .
An estimate of PET was obtained
from simple geometric considerations of the vertical distribution of
target areas (tree tops) and crown areas. First we assumed that target
and crown areas were distributed between a height of 10.2 and 26.2 m
(range = 16.0 m). These limits are the observed minimum and maximum laser
canopy height in the 36 field plots. Actual limits obtained from ground
data would be 5.5 to 31.7 m. Next, we posited a symmetrical bell-shaped
distribution of target and canopy areas. Magnussen and Boudewyn
(1998) found support for a beta distribution with parameters ± (=
2.1) and ² (= 2.1). Finally, we assumed that the projected crown
profiles were circular with a radius (CR) determined by the distance to
the tree top (DT) as in CR = DT0.421 (Magnussen and Boudewyn
1998). By computing the ratio of the expected target area to the expected
crown area in each of a series of canopy segments we essentially estimate
the probability that a target area is exposed within a segment. Canopy
segments were defined as a horizontal canopy slice with a width of 0.29 m
equal to the average height increment . Note the independence of view
angle: the area multipliers for target and crown areas due to off-nadir
view angles are identical and cancel when the taking ratios. Summing
these probabilities for all canopy segments yields the final
probability. In short:
(2)
where the symbols denotes the largest integer smaller than k and the smallest integer larger than
k. pdf(Beta( ± ,²),(x-10.2)/14.0) is the probability
density function of a beta distribution with parameters ± ( = 2.1) and ²
( = 2.1) evaluated at an interior point x of the interval
[10.2; 26.2]. The last term in (2) is a division by the total probability
of a target area.
2.10 Extracting the topmost canopy heights (CH)
To ensure a spatially balanced sample of the topmost canopy heights a double
stratified sampling scheme was applied. First, each plot was subdivided
into 
sections or strips along
an East-West direction, and then sectioned along a North-South
direction. Subsequently, for each of the two sectioning directions the
topmost laser canopy height in each section was selected as a proxy
height of dominant and codominant tree heights. Thus two samples of the
topmost 
canopy heights were
generated from each plot. The two samples, although not independent of
each other, are treated as simple replicates in the statistical
analyses.
Fitting an extreme value distribution to the topmost canopy
heights
Statistical inference for the selected topmost
canopy heights is limited by small sample sizes (here about 5 per plot
and direction of sectioning), the data (unequal number of canopy laser
heights in each section), and the selection process (the topmost). More
robust and extended inferences are possible if we can obtain the expected
distribution of the selected canopy heights. According to extreme value
theory (Smith 1986), when a sample of n canopy heights are independent and identically
distributed and there exist and such that the largest canopy height in the sample (CH1:n
) when scaled to converges in distribution to a
random variable with a non-degenerate cdf, and that cdf is one of three
types (Frechet, Weibull, or Gumbel). The convergence occurs as . In
other words, if we can derive estimates of the two parameters
(a,b) we would have a proxy distribution of TH. For convenience
and ease of estimation the Gumbel model was chosen to quantify the
distribution of extreme canopy heights.
Estimating the a and b parameters in the Gumbel
distribution
The maximum canopy height in a sample of n canopy heights
(CH1:n ) is assumed to have the following Gumbel
(G) probability density distribution (pdf) with parameters
a and b (Smith 1986):
From this proposition it follows that plot specific
estimates of the Gumbel parameters a and b based on
observations of the largest canopy height within each of sections in a plot allow us to
predict the pdf of heights of dominant and codominant trees. The
mean tree height and other descriptive statistics of dominant and
codominant trees are derived from this pdf by conventional
calculus (Johnson et al. 1994).
Plot specific maximum likelihood estimates of a and b
were obtained by maximizing the following likelihood (Strand and Boes
1998):

where subscript refers to plots ( ) and

is the total number of
included laser canopy heights in plot and is the number of included laser canopy heights in the
kth section of plot. As a rule we excluded the lower half of the
laser canopy heights from each plot section. This exclusion of the lower
half had virtually no impact on the estimated parameters but improved the
computational aspects considerably, especially convergence. Whenever fell
below three, the number of plot sections were lowered by one, and so on
until all sections contained at least three canopy heights. For these
plots n in (3) was lowered accordingly. Maximization of (4) was
done with a Newton-Raphson algorithm (McCulloch 1997).
Table 1 summarizes the ground-based maximum likelihood
results. CASI-based results were similar (not shown). All estimates of
a and b were more than 3 times as large as their estimated
standard deviations. The correlation between estimates of a and
b was strong and negative (-0.80). Estimates from the East-West
and the North-South sectioning were, in general, within 10% of each
other. Occassionally (four plots), when the spatial distribution of
canopy heights were very uneven, larger discrepancies emerged. For each
plot we used the simple average of the East-West and the North-South
estimates as the parameters of a plot specific Gumbel pdf of
presumed height of dominant and codominant trees.
Figure 1 shows the laser canopy heights in a randomly
chosen plot (no. 10) by East-West and North-South partitioning of the
plot. In this particular example only the topmost 4 canopy heights in
each section were considered for fitting. Figure 2 provides further
details on the estimated Gumbel pdf and the selected topmost laser canopy
heights.
Given estimates of ai and
bi, the predicted mean height of dominant and
codominant trees in the ith plot was:
(5) 
where is mean number of canopy heights used for estimation in plot
. The divisor in (5) is the total probability integral of the
Gumbel pdf. Integration limits were arbitrarily set to 0 and 40
m. Variance estimates of were computed by replacing the integrand h by .
3. Results and discussion
Ground- and CASI-based estimates of the number
of ‘visible’ trees (
) showed
considerable divergence. The minimum, mean, and median CASI-based counts
were about 50% above the estimates derived from purely geometric
considerations. This is hardly surprising because the geometric approach
treated the crowns and tree tops as solids and implicitly assumed trees
to be randomly distributed when in reality tree crowns are less than
opaque (Carlson and Ripley 1997), and trees in this trial tend to be
spaced more regularly than random ( , based on a Chi-Square goodness-of-fit statistics for the
quadrat method of testing of complete spatial randomness, Cressie
1991). Number of quadrats per plot was 20. Magnussen and Boudewyn
(1998) argued for a crown transparency of 50% (that is only half the
crown projection area is occupied by foliage, branches and stems) which
would help narrow the gap between the two estimates of . Less divergence emerged in the
expected number of tree top hits . Here the ground-based estimates surpassed the CASI-based
estimates by about 20%. On average we expect 4.7 tree top hits given the
ground-based approach and 5.8 given the CASI-based counts. Given
intrinsic differences in spatial resolution and hence definitions of
target area, the concordance is fortuitous.
Estimated average dominant and codominant tree height
derived from the expected number of tree tops hit and plot specific
Gumbel pdf’s were generally within one meter of estimated Lorey’s
heights (Figure 3). Simple means of the topmost canopy height from each plot were equally close to
their ground counterparts (not shown). Ground-based estimates were
slightly closer to Lorey’s heights than the CASI-based estimates. The
average difference of 13 cm was, however, not statistically significant
(
). In six plots the
discrepancy between estimated and actual height exceeds 3 m. Previous
analyses (Magnussen and Boudewyn 1998) made it clear that the six plots
could reasonably be considered as outliers. The number of laser canopy
hits were in all six instances well below average and spatially clustered
towards one end of the respective plots.
On average, Lorey’s mean height of 22.4 m exceeded the
ground-based estimate by 60 cm. A simple two-sided t-test did not reject
the hypothesis of no difference ( . The corresponding mean absolute deviation (MAD) was 1.34
m. Parallel CASI-based results were 73 cm ( ) with
MAD at 1.39 m and a correlation of 0.62. Standard deviations of the
recovered heights, as obtained from the fitted Gumbel pdf’s varied from
0.7 to 1.6 m (mean 1.0m). Thus most (30 of 36) estimated plot means of
dominant and codominant tree height had an estimated confidence interval
that included Lorey’s height. In contrast, mean laser canopy heights,
were, on average 4.3 m shorter than Lorey’s height ( and
MAD averaged 4.3m
A one-tree difference in the mean number of
expected hits between the ground- and CASI-based method explicates the
consistent off-set of about 20 cm in the point estimates. The off-set is
consistent with the differential selection intensity (Magnussen 1999). In
the CASI-based approach 1 laser canopy height were selected out of an
average of 8 while 1 of 10 were selected in the ground-based
approach. Assuming that heights of dominant and codominant trees are
nearly normally distributed the stated difference in selection intensity
alone accounts for 85% of the offset (see Falconer 1981 for details on
the effect of selection differential on expected means of selected
items).
Part of the persistent downward bias in our recovered
tree heights may actually be explicated by the selection procedure of
the topmost laser canopy heights in each plot. Our partitioning of each
plot and selection of only the topmost in each section inadvertently
yields lower heights than a selection of the topmost
canopy heights within the entire plot. Given the estimated standard
deviations of recovered height we expect a priori a downward bias of
about 20 cm. However, we rationalize that the spatial partitioning is
consistent with the averaging over all trees in a plot.
4. Conclusions
For laser scanning of forest canopies to become
a realistic forest inventory alternative to a direct
(traditional) measurement of tree heights it is key that easy-to-use
robust methods for extracting tree heights from canopy heights become
available (Magnussen and Boudewyn 1998, N¶sset 1997, Nilsson 1996). The
typical significant discrepancy between canopy heights and tree heights
prevents the direct use of canopy heights as a proxy for tree
height. Site index and volume estimates based on mean canopy heights
would be seriously downward biased. Results from this study confirmed
that easy-to-use methods transform laser canopy heights into good
approximations of desired tree heights.
Magnussen et al. (1999) showed that data processing
based on double-blind deconvolution is capable of removing the average
bias between laser-based and ground-based estimates of tree heights,
albeit at a higher price. Additional costs are to be paid for analysis
and computing.
Reasonable approximations of tree height have been
obtained from laser canopy data by selecting only a few topmost canopy
heights from within a plot (Tickle et al. 1998, Næsset 1997, Nilsson
1996). Local experiences have guided the establishment of sampling
guidelines and the results indicate that estimated tree heights are
typically within 0.5 to 2 m of the ground-based results when trees are
between 12 and 20 m tall. Magnussen and Boudewyn (1998) found support for
this practical approach in a thinning trial with Douglas-fir by
demonstrating that the vertical distribution of canopy heights obtained
from airborne laser scanning closely resembled the vertical distribution
of foliage surface. Thus, selecting a quantile of the canopy heights that
matches in probability the quantile of foliage surface area corresponding
to a desired tree height would produce unbiased estimates of tree
height.
A model-based approach to estimate tree heights
directly from canopy heights has the advantage of establishing a
framework for optimization and statistical inference. Derivation of tree
heights from canopy heights without auxiliary information is feasible
(Magnussen et al. 1999) but the necessary deconvolution algorithms are
computationally intensive.
Auxiliary ground information or information from
other remotely sensed images such as CASI can greatly reduce the
computational complexities of recovering tree heights from laser canopy
heights. A pivotal parameter to be gained from the auxiliary information
is the number of tree tops and size of tree crowns. Pragmatism must
prevail in the definition of ‘tree top’. A reasonable definition should
be possible in most circumstances.
Once an estimate of this number has been obtained, it
is straightforward to either use this number of topmost canopy heights
from each plot as proxies for tree heights or, as demonstrated, as input
to fit a distribution model of extreme values. The difference is bound to
be inconsequential. Fitting a distribution (say Gumbel) offers the
additional advantage of sufficient statistics of the estimated
quantities.
5. Acknowledgments
The CASI imagery was acquired under a Forest Renewal of British Columbia supported project entitled "Development of Certified Forestry Applications Using Compact Airborne Spectrographic Imager (CASI) Data", a co-operation between MacMillan Bloedel Ltd., ITRES Research Ltd., and the Canadian Forest Service (Pacific Forestry Centre).
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Table 1. Summary of ground-based maximum likelihood estimation
of the a and b parameters of the Gumbel pdf in 36 plots . "East-West”,
and "North-South” refer to cardinal direction of plot sectioning (see
text for details). Std. dev. is the among-plot standard deviation of a
parameter. RMSE is the root mean square error of a maximum likelihood
estimate as obtained from the inverse of minus the Fisher information
matrix (Gallant 1987).
|
Parameter |
Mean |
Minimum |
Maximum |
Std. dev. |
|

(East-West)
|
18.4 |
8.1 |
22.9 |
3.0 |
|

(North-South)
|
18.6 |
7.6 |
24.0 |
3.0 |
|
RMSE (
, East-West) |
0.40 |
0.21 |
0.86 |
0.12 |
|
RMSE (
, North-South) |
0.41 |
0.18 |
0.62 |
0.10 |
|

(East-West)
|
1.9 |
1.3 |
3.0 |
0.5 |
|

(North-South)
|
1.9 |
0.9 |
3.0 |
0.5 |
|
RMSE (
, East-West) |
0.17 |
0.08 |
0.46 |
0.07 |
|
RMSE (
, North-South) |
0.18 |
0.06 |
0.32 |
0.06 |
|
Sections (East-West) |
4.6 |
2 |
6 |
0.8 |
|
Sections (North-South) |
4.5 |
3 |
3 |
0.6 |
|
ni (East-West) |
4.2 |
3 |
11 |
1.4 |
|
ni (North-South) |
4.0 |
3 |
8 |
1.3 |
|

East-West
|
-46.1 |
-299.4 |
-8.3 |
158.9 |
|

North-South
|
-19.1 |
-34.1 |
-7.8 |
5.0 |
Table 2 . Summary of stem counts from CASI
images and a ground-based tally of 36 square 20 x 20 m plots.
|

a
laser |

ground |
b
ground
|
c
CASI |

ground |

CASI |
|
Min |
29 |
32 |
12.2 |
30 |
2.1 |
3.4 |
|
Median |
44 |
63.5 |
24.2 |
42 |
4.5 |
5.6 |
|
Mean |
48.2 |
66.0 |
25.1 |
42.2 |
4.7 |
5.8 |
|
Max |
71 |
158 |
60.2 |
55 |
8.4 |
9.9 |
|
Std. dev. |
11.4 |
31.8 |
12.1 |
6.3 |
1.6 |
1.5 |
Number of laser canopy hits per plot
Estimated number of visible tree tops per plot ( , see (2))
Tree count by local maxima procedures on CASI images (see text)
Figure 1. Canopy heights (CH) in m by plot section and maximum
likelihood estimates of the two Gumbel pdf parameters a and
b. Plot size = 20 x 20 m.
á
North-South partitioning (4 x 20m).
n
East-West Partitioning (20 x 4m).

Figure 2 . Histogram of tree height (HT, light
shaded bars) and 10 topmost canopy heights (darker shaded bars) in 5
East-West and 5 North-South 4 x 20 m plot sections in plot no. 10. The
fitted Gumbel pdf is superimposed. Vertical lines are mean of the Gumbel
extreme value distribution (full line) and Lorey’s height as per
ground-based measurements.
Figure 3. Estimated plot mean tree height of
dominant and codominant trees and canopy mean height versus estimates of
Lorey’s height (HT). LHT= laser-based height estimate.
ƒ
Estimates derived from ground-based tree counts
I
Estimates derived from CASI-based tree counts
n
Mean laser canopy height
Error bars indicate the interval of the estimate ±
one standard deviation.
|